diff --git a/README.md b/README.md
index 0b0b3adc..1e84692c 100644
--- a/README.md
+++ b/README.md
@@ -28,7 +28,7 @@ The `traccuracy` library has three main components: loaders, matchers, and metri
 Loaders load tracking graphs from other formats, such as the CTC format, into a [TrackingGraph](https://traccuracy.readthedocs.io/en/latest/autoapi/traccuracy/index.html#traccuracy.TrackingGraph) object.
 A TrackingGraph is a spatiotemporal graph.
 Nodes represent a single cell in a given time point, and are annotated with a time and a location.
-Edges point from a node representing a cell in time point `t` to the same cell or its daughter in `t+1`.
+Edges point forward in time from a node representing a cell in time point `t` to the same cell or its daughter in frame `t+1` (or beyond, to represent gap-closing).
 To load TrackingGraphs from a custom format, you will likely need to implement a loader: see
 documentation [here](https://traccuracy.readthedocs.io/en/latest/autoapi/traccuracy/loaders/index.html#module-traccuracy.loaders) for more information.
 
@@ -55,4 +55,7 @@ pipelines, [documented here](https://traccuracy.readthedocs.io/en/latest/cli.htm
 : A single non-dividing cell tracked over time. In graph terms, this is the connected component of a track between divisions (daughter to next parent). Tracklets can also start or end with a non-dividing cell at the beginning and end of the captured time or if the track leaves the field of view.
 
 **Track**
-: A single cell and all of its progeny. In graph terms, a connected component including divisions.
\ No newline at end of file
+: A single cell and all of its progeny. In graph terms, a connected component including divisions.
+
+**Gap-Closing**
+: Also known as *frame-skipping*, these are edges that connect non-consecutive frames to signify a cell being occluded or missing for some frames, before the track continues.
\ No newline at end of file
diff --git a/src/traccuracy/_tracking_graph.py b/src/traccuracy/_tracking_graph.py
index 082e0ac3..8f21bc62 100644
--- a/src/traccuracy/_tracking_graph.py
+++ b/src/traccuracy/_tracking_graph.py
@@ -81,6 +81,9 @@ class TrackingGraph:
     location (defaults to 'x' and 'y'). As in networkx, every cell must have a unique id, but these
     can be of any (hashable) type.
 
+    Edges typically connect nodes across consecutive frames, but gap closing or frame
+    skipping edges are valid, which connect nodes in frame t to nodes in frames beyond t+1.
+
     We provide common functions for accessing parts of the track graph, for example
     all nodes in a certain frame, or all previous or next edges for a given node.
     Additional functionality can be accessed by querying the stored networkx graph
diff --git a/tests/loaders/test_ctc.py b/tests/loaders/test_ctc.py
index 38dcc06f..a621fa4d 100644
--- a/tests/loaders/test_ctc.py
+++ b/tests/loaders/test_ctc.py
@@ -66,6 +66,23 @@ def test_ctc_single_nodes():
     TrackingGraph(G)
 
 
+def test_ctc_with_gap_closing():
+    data = [
+        {"Cell_ID": 1, "Start": 0, "End": 1, "Parent_ID": 0},
+        {"Cell_ID": 2, "Start": 0, "End": 1, "Parent_ID": 0},
+        # Connecting frame 1 to frame 3
+        {"Cell_ID": 3, "Start": 3, "End": 5, "Parent_ID": 1},
+        # Connecting frame 1 to frame 6
+        {"Cell_ID": 4, "Start": 6, "End": 8, "Parent_ID": 2},
+    ]
+    df = pd.DataFrame(data)
+    G = _ctc.ctc_to_graph(
+        df, pd.DataFrame({"segmentation_id": [], "x": [], "y": [], "z": [], "t": []})
+    )
+    assert G.has_edge("1_1", "3_3")
+    assert G.has_edge("2_1", "4_6")
+
+
 def test_load_data():
     test_dir = os.path.abspath(__file__)
     data_dir = os.path.abspath(
diff --git a/tests/metrics/test_ctc_metrics.py b/tests/metrics/test_ctc_metrics.py
index bd92fc95..e8885b8a 100644
--- a/tests/metrics/test_ctc_metrics.py
+++ b/tests/metrics/test_ctc_metrics.py
@@ -1,7 +1,9 @@
+from traccuracy._tracking_graph import EdgeAttr, NodeAttr, TrackingGraph
+from traccuracy.matchers._base import Matched
 from traccuracy.matchers._ctc import CTCMatcher
 from traccuracy.metrics._ctc import CTCMetrics
 
-from tests.test_utils import get_movie_with_graph
+from tests.test_utils import get_gap_close_graphs, get_movie_with_graph
 
 
 def test_compute_mapping():
@@ -17,3 +19,19 @@ def test_compute_mapping():
     assert "DET" in results
     assert results["TRA"] == 1
     assert results["DET"] == 1
+
+
+def test_compute_metrics_gap_close():
+    g_gt, g_pred, mapper = get_gap_close_graphs()
+    matched = Matched(
+        gt_graph=TrackingGraph(g_gt), pred_graph=TrackingGraph(g_pred), mapping=mapper
+    )
+    CTCMetrics().compute(matched)
+
+    # check that missing gap closing edge is false negative
+    assert g_gt.edges[("1_1", "2_3")][EdgeAttr.FALSE_NEG]
+    # check that "extra" node is FP
+    assert g_pred.nodes["1_2"][NodeAttr.FALSE_POS]
+    # check that correct edge is not annotated with errors
+    for error_attr in [EdgeAttr.FALSE_POS, EdgeAttr.WRONG_SEMANTIC]:
+        assert not g_pred.edges[("2_6", "4_10")][error_attr]
diff --git a/tests/metrics/test_track_overlap_metrics.py b/tests/metrics/test_track_overlap_metrics.py
index f70a9304..8a2da45e 100644
--- a/tests/metrics/test_track_overlap_metrics.py
+++ b/tests/metrics/test_track_overlap_metrics.py
@@ -6,6 +6,8 @@
 from traccuracy.matchers import Matched
 from traccuracy.metrics._track_overlap import TrackOverlapMetrics, _mapping_to_dict
 
+from tests.test_utils import get_gap_close_graphs
+
 
 def add_frame(tree):
     attrs = {}
@@ -183,6 +185,19 @@ def test_track_overlap_metrics(data, inverse) -> None:
     assert results == expected, f"{data['name']} failed without division edges"
 
 
+def test_track_overlap_gap_close():
+    g_gt, g_pred, mapping = get_gap_close_graphs()
+    matched = Matched(
+        TrackingGraph(g_gt),
+        TrackingGraph(g_pred),
+        mapping,
+    )
+    metric = TrackOverlapMetrics()
+    results = metric.compute(matched)
+    assert results["track_purity"] == 7 / 9
+    assert results["target_effectiveness"] == 7 / 8
+
+
 def test_mapping_to_dict():
     mapping = [("1", "2"), ("2", "3"), ("1", "3"), ("2", "3")]
     mapping_dict = _mapping_to_dict(mapping)
diff --git a/tests/test_utils.py b/tests/test_utils.py
index 1dbf6d7c..9d98cdb9 100644
--- a/tests/test_utils.py
+++ b/tests/test_utils.py
@@ -148,3 +148,124 @@ def get_division_graphs():
     mapper = [("1_0", "1_0"), ("1_1", "1_1"), ("2_4", "2_4"), ("3_4", "3_4")]
 
     return G1, G2, mapper
+
+
+def get_gap_close_graphs():
+    """
+    G1
+                                      3_5 -- 3_6 -- -- -- 5_10
+    1_0 -- 1_1 -- -- -- 2_3 -- 2_4 -<
+                                      4_5 -- 4_6
+    G2
+                                      2_5 -- 2_6 -- -- -- 4_10
+    1_0 -- 1_1 -- 1_2 -- 1_3 -- 1_4 -<
+                                      3_5 -- 3_6
+    """
+    G1 = nx.DiGraph()
+    G1.add_edge("1_0", "1_1")
+    # gap closing edge
+    G1.add_edge("1_1", "2_3")
+    G1.add_edge("2_3", "2_4")
+    # Divide to generate 3 lineage
+    G1.add_edge("2_4", "3_5")
+    G1.add_edge("3_5", "3_6")
+    # gap closing edge
+    G1.add_edge("3_6", "5_10")
+    # Divide to generate 4 lineage
+    G1.add_edge("2_4", "4_5")
+    G1.add_edge("4_5", "4_6")
+
+    attrs = {}
+    for node in G1.nodes:
+        attrs[node] = {"t": int(node[-1:]), "x": 0, "y": 0}
+    nx.set_node_attributes(G1, attrs)
+
+    G2 = nx.DiGraph()
+    G2.add_edge("1_0", "1_1")
+    # missing gap closing edge
+    G2.add_edge("1_1", "1_2")
+    G2.add_edge("1_2", "1_3")
+    G2.add_edge("1_3", "1_4")
+    # Divide to generate 2 lineage
+    G2.add_edge("1_4", "2_5")
+    G2.add_edge("2_5", "2_6")
+    # correct gap closing edge
+    G2.add_edge("2_6", "4_10")
+    # Divide to generate 3 lineage
+    G2.add_edge("1_4", "3_5")
+    G2.add_edge("3_5", "3_6")
+
+    attrs = {}
+    for node in G2.nodes:
+        attrs[node] = {"t": int(node[-1:]), "x": 0, "y": 0}
+    nx.set_node_attributes(G2, attrs)
+
+    # G1, G2 mapper
+    mapper = [
+        ("1_0", "1_0"),
+        ("1_1", "1_1"),
+        ("2_3", "1_3"),
+        ("2_4", "1_4"),
+        ("3_5", "2_5"),
+        ("3_6", "2_6"),
+        ("5_10", "4_10"),
+        ("4_5", "3_5"),
+        ("4_6", "3_6"),
+    ]
+
+    return G1, G2, mapper
+
+
+def get_division_gap_close_graphs():
+    """
+    G1
+                   -- -- 2_3 -- 2_4
+    1_0 -- 1_1 -<
+                  3_2 -- 3_3 -- 3_4
+    G2
+                  2_2 -- 2_3 -- 2_4
+    1_0 -- 1_1 -<
+                  3_2  -- -- -- 4_4
+    """
+
+    G1 = nx.DiGraph()
+    G1.add_edge("1_0", "1_1")
+    # gap division
+    G1.add_edge("1_1", "2_3")
+    G1.add_edge("2_3", "2_4")
+    # divide into 3 lineage
+    G1.add_edge("1_1", "3_2")
+    G1.add_edge("3_2", "3_3")
+    G1.add_edge("3_3", "3_4")
+
+    attrs = {}
+    for node in G1.nodes:
+        attrs[node] = {"t": int(node[-1:]), "x": 0, "y": 0}
+    nx.set_node_attributes(G1, attrs)
+
+    G2 = nx.DiGraph()
+    G2.add_edge("1_0", "1_1")
+    # Divide to generate 2 lineage
+    G2.add_edge("1_1", "2_2")
+    G2.add_edge("2_2", "2_3")
+    G2.add_edge("2_3", "2_4")
+    # Divide to generate 3 lineage
+    G2.add_edge("1_1", "3_2")
+    # incorrect gap closing edge
+    G2.add_edge("3_2", "4_4")
+
+    attrs = {}
+    for node in G2.nodes:
+        attrs[node] = {"t": int(node[-1:]), "x": 0, "y": 0}
+    nx.set_node_attributes(G2, attrs)
+
+    mapper = [
+        ("1_0", "1_0"),
+        ("1_1", "1_1"),
+        ("2_3", "2_3"),
+        ("2_4", "2_4"),
+        ("3_2", "3_2"),
+        ("3_4", "4_4"),
+    ]
+
+    return G1, G2, mapper
diff --git a/tests/track_errors/test_divisions.py b/tests/track_errors/test_divisions.py
index 6538e644..8822c75e 100644
--- a/tests/track_errors/test_divisions.py
+++ b/tests/track_errors/test_divisions.py
@@ -11,7 +11,7 @@
     _get_succ_by_t,
 )
 
-from tests.test_utils import get_division_graphs
+from tests.test_utils import get_division_gap_close_graphs, get_division_graphs
 
 
 @pytest.fixture
@@ -220,3 +220,30 @@ def test_evaluate_division_events():
     results = _evaluate_division_events(matched_data, frame_buffer=frame_buffer)
 
     assert np.all([isinstance(k, int) for k in results.keys()])
+
+
+def test_gap_close_divisions():
+    g_gt, g_pred, mapper = get_division_gap_close_graphs()
+    matched_data = Matched(TrackingGraph(g_gt), TrackingGraph(g_pred), mapper)
+    _classify_divisions(matched_data)
+
+    # missing gap close div edge so FN DIV
+    assert g_gt.nodes["1_1"][NodeAttr.FN_DIV]
+
+    # fix division, assert it's identified correctly
+    g_pred.remove_node("2_2")
+    g_pred.add_edge("1_1", "2_3")
+    # mapper doesn't need to change as removed node was always missing
+    matched_data = Matched(TrackingGraph(g_gt), TrackingGraph(g_pred), mapper)
+    _classify_divisions(matched_data)
+    assert g_gt.nodes["1_1"][NodeAttr.TP_DIV]
+    assert g_pred.nodes["1_1"][NodeAttr.TP_DIV]
+
+    g_gt, g_pred, mapper = get_division_gap_close_graphs()
+    # remove gt division
+    g_gt.remove_edge("1_1", "2_3")
+    g_gt.remove_edge("1_1", "3_2")
+    matched_data = Matched(TrackingGraph(g_gt), TrackingGraph(g_pred), mapper)
+    _classify_divisions(matched_data)
+    # assert fp division classified correctly
+    assert g_pred.nodes["1_1"][NodeAttr.FP_DIV]